Referential and general calls in primate semantics

In recent years, the methods of formal semantics and pragmatics have been fruitfully applied to analysis primate communication systems. Most analyses therein appeal a division labor between which has following three features: (F1) calls are given referential meanings (they provide information about world rather than just an action be taken), (F2) some general meaning, (F3) in context enriched by competition with more informative calls, along lines scalar implicatures. this paper, we develop highly simplified models independently assess conditions under such features would emerge. After identifying sufficient condition for (F1), find range not evolutionarily stable, discuss consequences both modeling empirical work.